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Sandra Wilsher BSc (Hons) PhD
Sandra spent a decade working in the British horse industry before undertaking a Biology degree, graduating three years later with a First Class Honours degree and the Dean’s Prize for the Best Science Student. She joined Professor Twink Allen’s Equine Fertility Unit (EFU) in worked until its closure in 2007. She has published over 35 papers in peerreviewed journals on fetal and maternal factors that govern the development and functions of the equine placenta, equine embryo transfer, and reproductive efficiency and wastage in the Thoroughbred racing industry. In addition, she has contributed to veterinary textbooks and presented at numerous conferences and workshops both in the UK and abroad to veterinary, scientific and lay audiences. While at the EFU she developed the Wilsher Equine Embryo Transfer Forceps which increased the non- surgically ET rates at the EFU significantly and are now produced commercially. Shortly after the EFU closed she joined Professor Allen at the Paul Mellon Laboratory of Equine Reproduction where she completed her PhD by Published Works and where she continues to work both on equine reproduction and, more recently, placentation and other aspects of reproduction in the giraffe, zebra and wildebeest. In addition, she advises and assists the two local equine veterinary practices with their embryo transfer
Bibliography Refereed papers

1. Wilsher S, Ball M and Allen WR (1999) The influence of maternal size on placental area
and foal birthweights in the mare. Pferdeheilkunde 15 599-602
2. Allen WR, Wilsher S, Turnbull C, Stewart F, Ousey J, Rossdale PD and Fowden A (2002)
Influence of maternal size on placental, fetal and postnatal growth in the horse: I. Development in utero. Reproduction 123 445-453
3. Allen WR, Wilsher S, Stewart F, Ousey J and Fowden A (2002) The influence of maternal
size on placental, fetal and postnatal growth in the horse: II. Endocrinology of
pregnancy. Journal of Endocrinology 172 237-246
4. Wilsher S and Allen WR (2002) The influences of maternal size, age and parity on
placental and fetal development in the horse. Theriogenology 58 833-835
5. Wilsher S and Allen WR (2003) The effects of maternal age and parity on placental and
fetal development in the mare. Equine Veterinary Journal 35 476-483
6. Allen WR, Wilsher S, Tiplady C and Butterfield RM (2004) The influence of maternal size
on pre- and postnatal growth in the horse: III Postnatal growth. Reproduction 127 67-77
7. Wilsher S and Allen WR (2004) An improved method for non-surgical embryo transfer in
the mare. Equine Veterinary Education 16 39-44
8. Wilsher S, Allen WR and Wood JLN (2006) Factors associated with failure of
Thoroughbred horses to train and race. Equine Veterinary Journal 38 113-118
9. Xihe Li, Qin Y, Wilsher S and Allen WR (2006) Centrosome changes during meiosis in
horse oocytes and first embryonic cell cycle organization following parthenogenesis,
fertilization and nuclear transfer. Reproduction 131 661-667
10. Abd-Elnaeim MMM, Leiser R, Wilsher S and Allen WR (2006) Structural and
haemovascular aspects of placental growth throughout gestation in young and aged mares. Placenta 27 1103-1113
11. Wilsher S, Kölling M and Allen WR (2006) Meclofenamic acid extends donor-recipient
asynchrony in equine embryo transfer. Equine Veterinary Journal 38 428-432
12. Wilsher S and Allen WR (2006) Effects of a Streptococcus equi infection mediated
nutritional insult during mid-gestation in Thoroughbred mares. I. Placental and fetal development. Equine Veterinary Journal 38 549-557
13. Allen WR, Wilsher S, Morris L, Crowhurst JS, Hillyer MH, Neal HN (2006) Laparoscopic
application of PGE2 to re-establish oviducal patency and fertility in infertile mares: a preliminary study Equine Veterinary Journal 38 454-459
14. Allen WR, Gower S and Wilsher S (2007) Immunohistochemical localisation of vascular
endothelial growth factor (VEGF) and its two receptors (Flt-I and KDR) in the endometrium and placenta of the mare during 15. Allen WR, Brown L, Wright M and Wilsher S (2007) Reproductive efficiency of Flatrace
and National Hunt Thoroughbred mares and stallions in England. Equine Veterinary Journal 39 438-445
16. Allen WR, Kölling M and Wilsher S (2007) An interesting case of early pregnancy loss in a
mare with persistent endometrial cups. Equine Veterinary Education 19 539-544
17. Ellenberger C, Wilsher S, Allen WR, Hoffman C, Kölling M, Bazer FW, Klug J, Schoon D and
Schoon HA (2008) Immunolocalisation of the uterine secretory proteins uterocalin, uteroferrin and uteroglobulin in the mare’s uterus and placenta throughout gestation. Theriogenology 70 746-757
18. Ousey JC, Fowden AL, Wilsher S and Allen WR (2008) The effects of maternal health and
body condition on the endocrine responses of neonatal foals. Equine Veterinary Journal 40 673-679
19. Abd-Elnaeim MM, Derar IR, Wilsher S, Allen R, Leiser R and Schuler G (2009)
Immunohistochemical localization of oestrogen receptors alpha and beta, progesterone receptor and aromatase in the equine placenta. Reproduction in Domestic Animals 44
20. Wilsher S and Allen WR (2009) Uterine influences on embryogenesis and early
placentation in the horse revealed by transfer of day-10 embryos to day-3 recipient mares. Reproduction 137 583-593
21. Ellenberger C, Muller K, Schoon HA, Wilsher S and Allen WR (2009) Histological and
immunohistochemical characterization of equine anovulatory haemorrhagic follicles (AHFs). Reproduction in Domestic Animals 44 395-405
22. Wilsher S, Lefranc A-C and Allen WR (2009) Postnatal oestrogen administration
stimulates precocious endometrial gland development in the horse. Equine Veterinary Journal 41 678-684
23. Wilsher S, Ousey JC and Allen WR (2009) Abnormal umbilical cord attachment sites in the
mare: a review illustrated by three case reports. Equine Veterinary Journal 41 930-939
24. Allen WR and Wilsher S (2009) A review of implantation and early pregnancy in the
mare. Placenta 30 1005-1015
25. Wilsher S, Clutton-Brock A and Allen WR (2010) Successful transfer of day-10 equine
embryos: Influence of donor-recipient asynchrony on embryo development. Reproduction 139 575-585
26. Veronesi MC, Villani M, Wilsher S, Contri A and Carluccio A (2010) A comparative study
of the term placenta in the donkey, pony and thoroughbred. Theriogenology 74 627-631
27. Wilsher S and Allen WR (2011) Intrauterine administration of plant oils inhibits luteolysis
in the mare. Equine Veterinary Journal 43 99-105
28. Wilsher S, Ousey J, Whitwell K, Crowhurst JS, Neal HN and Allen WR (2011) 3 types of
anomalous vasculature in the equine umbilical cord. Equine Veterinary Education 23
29. Wilsher S and Allen WR (2011) Factors influencing equine Chorionic Gonadotrophin
(eCG) production in the mare. Equine Veterinary Journal 43 430-438
30. Wilsher S, Gower S and Allen WR (2011) Immunohistochemical localisation of
progesterone and oestrogen receptors at the placental interface in mares during early pregnancy. Animal Reproduction Science 129 200-208
31. Wilsher S and Allen WR (2012) Factors influencing placental development and function in
the mare. Equine Veterinary Journal 44 Suppl 41 113-119
32. Allen WR and Wilsher S (2012) Persistent endometrial cups in the same mare in two
successive pregnancies. Equine Veterinary Education 24 247-250
33. Wilsher S, LeFranc A-C and Allen WR (2012) The effects of an advanced uterine
environment on embryonic survival in the mare. Equine Veterinary Journal 44 432–439
34. Allen WR and Wilsher S (2012) The influence of season, mare numbers and ejaculation
frequency on the fertility of Thoroughbred stallions. Equine Veterinary Journal 44 535-
541 doi: 10.1111/j.2042-3306.2011.00525.x 35. Wilsher S, Ousey J and Allen WR (2013) Observations on the placentae of 8
Thoroughbred foals born with flexural limb deformities. Equine Veterinary Education 25
84-95 doi: 10.1111/j.2042-3292.2012.00433.x 36. Antczak DF, De Mestre A, Wilsher S and Allen WR (2013) The Equine Endometrial Cup
Rction: A Fetomaternal Signal of Significance. Annual Review of Animal Biosciences Vol 1
8.1 – 8.24 doi: 1146/annurev-animal-031412-103703
37. Wilsher S, Standfield FJ, Greenwood RES, Trethowan PD, Anderson RA, Wooding FB and
Allen WR (2013) Ovarian and placental morphology and endocrine functions in the pregnant giraffe (Giraffa camelopardalis) Reproduction 145 541-554
38. Wilsher S, Gower S and Allen WR (2013) Persistence of an immunoreactive MUC1
protein at the feto-maternal interface throughout pregnancy in the mare. Reproduction, Fertility and Development 25 753-61. doi: 10.1071/RD12152
Book chapters and course notes
1. Wilsher S and Allen WR (2004) Factors controlling microcotyledon development and
placental efficiency in Thoroughbreds: Influences of mare age, parity and nutritional insults. In: Proceedings of a Workshop on Equine Placenta. Kentucky Agricultural Experiment Station, Lexington, Kentucky, SR-2004-1, pp58-64. 2. Wilsher S and Allen WR (2004) Factors influencing placental and fetal development in
the mare: important considerations for the selection of embryo recipients. In: Emerging Equine Science. Eds: J Alliston, S Chadd, A Ede, A Hemmings, J Hyslop, A Longland, H Moreton and Moore-Colyer M. Nottingham University Press, Nottingham, UK. pp87-89 3. Wilsher S and Allen WR (2009) Survey of fertility statistics of horse bred for Flat and
National Hunt racing with particular reference to equine pregnancy failure. In: Rossdale and Partners Equine Pregnancy Failure Course. Ed S. Ricketts, Lifelearn Ltd, Newmarket 4. Wilsher S and Allen WR (2009) Does lymphocyte immunisation have a role in the
prevention of equine early pregnancy failure? In: Rossdale and Partners Equine Pregnancy Failure Course. Ed S. Ricketts, Lifelearn Ltd, Newmarket UK. pp 53-61 5. Ousey JC, Wilsher S and Allen WR (2009) A survey of placentae associated with abnormal
foals. In: Rossdale and Partners Equine Pregnancy Failure Course. Ed S. Ricketts, Lifelearn 6. Allen WR and Wilsher S (2009) Veterinary aspects of Thoroughbred breeding: Past
successes and future prospects. The Havemeyer Foundation (Information pamphlet for 7. Wilsher S (2010) Advances in mare reproductive science and neonatal health. BSc
Module 38. The Open College of Equine Studies 8. Wilsher S and Allen WR (2011) Development and morphology of the placenta. In: Equine
Reproduction 2nd Edn. Eds. AO McKinnon, EL Squires, WE Vaala and D Varner. Vol 2: Ch 9. Allen WR, Gower S and Wilsher S (2011) Fetal membrane differentiation, implantation
and early placentation in the mare. In: Equine Reproduction 2nd Edn. Eds. AO McKinnon, EL Squires, WE Vaala and D Varner. Vol 2: Ch 226 pp 2187-2199 Wiley Blackwell 10. Allen WR and Wilsher S (2011) The influence of mare numbers and ejaculation frequency
on the fertility of Thoroughbred stallions. In: Rossdale and Partners Stallion Sub-fertility Course. Ed S. Ricketts, pp 1-3, Whorl Publishing Ltd, Newmarket UK 11. Allen WR and Wilsher S (2011) Equine embryo transfer and Artificial insemination.
Sharjah Equine Hospital, Sharjah, UAE. (Course booklet for a theoretical and practical Short Course for horse breeders and equine veterinary surgeons) 12. Wilsher S (2012) The use of intrauterine plant oil to supress oestrous behaviour in fillies.
Newmarket Equine Hospital: Advances in Equine Reproduction Course Notes. 13. Wilsher S (2012) Examination of the term placenta – abnormalities and quirks.
Newmarket Equine Hospital: Advances in Equine Reproduction Course Notes 14. Wilsher S (2012) Setting up a successful embryo transfer programme: Recipient
asynchrony – what you can get away with. Newmarket Equine Hospital: Advances in 15. Wilsher S (2013) Equine embryo transfer: Do you know what you’re looking for? BEVA
16. Wilsher S (2013) The role of the placenta in fetal development. In: Equine Neonatology
Eds: P Morresey and B Waldridge. Wiley Blackwell (in press) Abstracts
1. Wilsher S and Allen WR (1999) Influence of maternal age on placental function in the
mare. Proceedings of the 50th Annual Meeting of the European Association Animal 2. Allen WR, Wilsher S, Barrett K, Stewart F, Fowden A, Ousey J and Rossdale PD (2000)
Effects of maternal size on pre- and postnatal development in the horse. In: Fetomaternal Control of Pregnancy. Eds TAE Stout and JF Wade. Havemeyer Foundation Monograph Series 2 49-50 (Abstr)
3. Wilsher S, Ball M and Allen WR (2000) The influence of maternal size, parity and age on
morphometric placental development and function in the mare. In: Fetomaternal Control of Pregnancy. Eds TAE Stout and JF Wade. Havemeyer Foundation Monograph Series 2 51-52 (Abstr)
4. Wilsher S and Allen WR (2000) Influence of maternal size, age and parity on placental
and fetal development in the horse. Journal of Reproduction and Fertility Abstract Series
25 14 (Abstr)
5. Morris L H-A, Tiplady C, Cook E, Wilsher S, Xihe L and Allen WR (2000) Hysteroscopic
insemination of mares with low numbers of frozen-thawed ejaculated and epididymal spermatozoa. Journal of Reproduction and Fertility Abstract Series 25 29 (Abstr)
6. Neal H, Morris LHA, Wilsher S and Allen WR (2000) Laparoscopic embryo transfer.
Proceedings of 39th British Veterinary Association Congress. Birmingham UK. 117 (Abstr) 7. Allen WR and Wilsher S (2002) Influence of maternal uterine size on pre- and postnatal
development of the foal. Reproduction in Domestic Animals 37 223 (Abstr)
8. Li X, Morris LH-A, Wilsher S and Allen WR (2003) Fetal development after
intracytoplasmic sperm injection (ICSI) in the horse. In: From Epididymis to Embryo. Eds LH-A Morris, L Foster and JF Wade. Havemeyer Foundation Monograph Series 6 75-76
9. Allen WR and Wilsher S (2003) Influence of maternal size on pre- and post natal growth
of the foal. In: Comparative Neonatology and Perinatology. Eds P Sibbons, L Foster and JF Wade. Havemeyer Foundation Monograph Series 8 49-50 (Abstr)
10. Wilsher S and Allen WR (2003) Influences of maternal and fetal genotypes on placental
and fetal development in the horse. In: Comparative Neonatology/Perinatology. Eds. P Sibbons, L Foster and JF Wade. Havemeyer Foundation Monograph Series 8 51-53 (Abstr)
11. Wilsher S and Allen W R (2003) The effects of maternal nutrition on placental and fetal
development in maiden Thoroughbred mares. In: Embryonic and Fetal Nutrition Eds. S
Wilsher and J Wade. Havemeyer Foundation Monograph Series 10 70-71 (Abstr)
12. Wilsher S and Allen WR (2003) A novel method for non-surgical embryo transfer in the
mare. Reproduction in Domestic Animals 38 320 (Abstr)
13. Xihe Li, A-C Lefranc, S Wilsher and W R Allen (2004) What is different about the horse
embryo? In: European Equine Gamete Group (EEGG). Eds J Müller, Z Müller and JF Wade. Havemeyer Foundation Monograph Series 13 62 (Abstr)
14. Wilsher S and Allen WR (2004) A new method for non-surgical embryo transfer in the
horse. In: European Equine Gamete Group (EEGG). Eds J Müller, Z Müller and JF Wade. Havemeyer Foundation Monograph Series 13 42-43 (Abstr)
15. Xihe L, Lefranc AC, Wilsher S and Allen WR (2004) The characteristics of horse
trophectoderm cells and their ability to support embryonic development following nuclear transfer. Proceedings of 15th International Conference on Animal Reproduction.
Abstracts
Vol 2 561 (Abstr)
16. Wilsher S and Allen WR (2004) The effects of maternal nutrition on placental
development in maiden mares. Proceedings of 15th International Conference on Animal Reproduction. Abstracts Vol 1 94 (Abstr)
17. Wilsher S and Allen WR (2005) A novel method for non-surgical embryo transfer in the
mare. In: Proceedings of the 6th International Symposium on Equine Embryo Transfer. Eds. M Alvarenga and JF Wade. Havemeyer Foundation Monograph Series 14 110-112
(Abstr)
18. Allen WR, Wilsher S, Kölling M and Lefranc A-C (2005) Luteal and endometrial
interdependence for early embryonic development in the mare. In: Maternal Recognition of Pregnancy in the Mare III. Eds TAE Stout and JF Wade. Havemeyer Foundation Monograph Series 16 3-4 (Abstr)
19. Wilsher S, Kölling M and Allen WR (2005) The use of meclofenamic acid to extend donorrecipient
asynchrony in equine embryo transfer. In: Maternal Recognition of Pregnancy in the Mare III. Eds TAE Stout and JF Wade. Havemeyer Foundation Monograph Series 16
20. Wilsher S and Allen WR (2005) Factors influencing placental and fetal growth.
Proceedings of 44th Congress British Equine Veterinary Association. 69 (Abstr) 21. Wilsher S, Kölling M and Allen WR (2005) The use of meclofenamic acid to extend donorrecipient
asynchrony in equine embryo transfer. Reproduction in Domestic Animals 40
344 (Abstr)
22. Wilsher S, Ousey J and Allen WR (2005) Gross and histological observations on placentae
from abnormal pregnancies. In: Proceedings of a Workshop on Comparative Placentology. Eds. P Sibbons and JF Wade. Havemeyer Foundation Monograph Series 17
23. Wilsher S, Allen WR and Wood JLN (2005) Success and wastage rates in Thoroughbred
flat racing in Britain and Ireland. In: Thoroughbred Racing and Breeding Seminar; Cheltenham 2005 75 (Abstr)
24. Wilsher S, Kölling M and Allen WR (2006) Meclofenamic acid extends donor-recipient
asynchrony in equine embryo transfer. In: International Equine Gamete Group. Eds. H Alm, H Torner and JF Wade. Havemeyer Foundation Monograph Series 18 56-57 (Abstr)
25. Wilsher S, Kölling M, Allen WR (2006) The effects of estrogen during early pregnancy in
the mare on serum progestagen and IFG-1 levels and embryonic growth. Animal Reproduction Science 94 381-382 (Abstr)
26. Allen WR, Wilsher S, Morris L, Crowhurst JS, Hillyer MH, Neal HN (2006) Reestablishment
of ovdicual patency and fertility in infertile mares. Animal Reproduction Science 94 242-243
27. Villani M, Wilsher S, Carluccio A, Contri A and Veronesi MC (2006) Stereology in the term
placenta of the donkey, Thoroughbred and pony: A comparative study. Animal Reproduction Science 94 411-412 (Abstr)
28. Ellenberger C, Kölling, Wilsher S, Allen WR, Bazer FW, Klug J, Schoon D and Schoon H-A.
(2006) Immunolocalisation of uterine secretory proteins in the mare during the oestrous cycle and pregnancy. Reproduction in Domestic Animals 41 308 (Abstr)
29. Wilsher S, Kölling M and Allen WR (2006) The effects of exogenous oestradiol on luteal
function and embryonic growth during early pregnancy in the mare. Reproduction in Domestic Animals 41 360 (Abstr)
30. Allen WR, Wilsher S, Morris L, Crowhurst JS, Hillyer MH and Neal HN (2006) Reestablishment
of oviducal patency and fertility in infertile mares. Reproduction in Domestic Animals 41 360 (Abstr)
31. Wilsher S and Allen WR (2007) Intrauterine administration of plant oils suppresses
luteolysis in the mare. Proceedings of the 5th Joint Meeting of the UK Fertility Societies. 32. Allen WR, Brown L, Wright M and Wilsher S (2007) Reproductive efficiency of the
Thoroughbred. Proceedings of the 5th Joint Meeting of the UK Fertility Societies. 74 33. Wilsher S and Allen WR (2007) Successful transfer of Day 10 horse embryos to
asynchronous recipient mares. Reproduction in Domestic Animals 42 74 (Abstr)
34. Allen WR and Wilsher S (2008) The equine endometrial gland: a bountiful milch-cow of
pregnancy in the mare. In: Second Workshop on Embryonic and Fetal Nutrition. Eds. S
Wilsher, WR Allen and JF Wade. Havemeyer Foundation Monograph Series 21 47-50
35. Wilsher S and Allen WR (2008) Effects of a Streptococcus equi infection-mediated
nutritional insult during mid-gestation on placental and fetal development in primiparous Thoroughbred fillies. In: Second Workshop on Embryonic and Fetal Nutrition. Eds. S
Wilsher, WR Allen and JF Wade. Havemeyer Foundation Monograph Series 21 47-50
36. Ousey J, Fowden AL, Wilsher S and Allen WR (2008) Effects of maternal nutrition state
during pregnancy on insulin secretion and sensitivity in neonatal foals. In: Second Workshop on Embryonic and Fetal Nutrition. Eds. S Wilsher, WR Allen and JF Wade.
Havemeyer Foundation Monograph Series 21 76-79 (Abstr)
37. Allen WR, Wilsher S, Morris L H-A, Crowhurst JS, Hillyer MH and Neal HN (2008)
of fertility in ageing mares by laparoscopic application of PGE2 to their oviducts. Pferdeheilkunde 24 117 (Abstr)
38. Wilsher S and Allen WR (2008) Maternal recognition of pregnancy in the mare: Inhibition
of luteolysis by intrauterine administration of plant oils. Pferdeheilkunde 24 118 (Abstr)
39. Allen WR, Brown L, Wright M and Wilsher S (2008) Reproductive efficiency of well
managed Thoroughbred mares and stallions. Pferdeheilkunde 24 125 (Abstr)
40. Wilsher S, Allen WR and Woods JLN (2008) Success and wastage rates in Thoroughbred
flatracing in Britain. Pferdeheilkunde 24 125-126 (Abstr)
41. Wilsher S, Clutton-Brock A and Allen WR (2008) Transfer of day 10 embryos to
asynchronous recipient mares. Proceedings of the Havemeyer 7th International Symposium on Equine Embryo Transfer. Ed: WR Allen. R&W Publications, Newmarket, UK 42. Wilsher S, Clutton-Brock A, Allen WR and Morris L.H-A. (2008) Transfer of embryos to
oestrous recipient mares treated with long-acting progesterone or altrenogest. Proceedings of the Havemeyer 7th International Symposium on Equine Embryo Transfer. Ed: WR Allen. R&W Publications, Newmarket, UK. 77-78 (Abstr) 43. Wilsher S, Clutton-Brock A and Allen WR (2009) Non-surgical transfer of horse embryos
to asynchronous recipient mares. Proceedings of the 25th Scientific Meeting of the European Embryo Transfer Association, Poznan, Poland. p. 274 (Abstr) 44. Wilsher S and Allen WR (2010) Inhibition of luteolysis in the mare by plant oils. Animal
Reproduction Science 121S S58-59 (Abstr)
45. Wilsher S and Allen WR (2009) Factors influencing gonadotrophin production by the
equine invasive trophoblast. Annual CTR Trophoblast Meeting, Cambridge (Abstr) 46. Allen WR, Gower S and Wilsher S (2010) An immunohistochemical study of equine
placentation. Animal Reproduction Science 121S S316-317 (Abstr)
47. Wilsher S, Clutton-Brock A and Allen WR (2012) Influence of exogenous progesterone on
survival of asynchronously transferred Day-10 horse embryos. Proceedings of the 8th International Symposium on Equine Embryo Transfer in Journal of Equine Veterinary Science 32 413 (Abstr)
48. Wilsher S (2013) The equine placenta – a vital clue to neonatal problems. Colloquium for
Equine Reproduction. Proceedings of the British Society of Animal Science A239 (Abstr) 49. WR Allen, Stansfield F and Wilsher S (2013) Placentation in the giraffe, zebra and
wildebeest. Annual CTR Trophoblast Meeting, Cambridge (Abstr) 50. Wilsher S (2013) What’s in the flush? Proceedings of 52ndCongress British Equine
Veterinary Association. (Abstr) Invited oral communications
1. 1st International Conference on Equine Reproductive Medicine, Leipzig, Germany. 2. Havemeyer Foundation Workshop on Fetomaternal Control of Pregnancy, Barbados, 3. Havemeyer Foundation Workshop on Comparative Neonatology and Perinatology, 4. Argentinean Symposium on Equine Reproduction, Tandil, Argentina. November, 2002 5. New Zealand Link Foundation Breeders Symposium, Agricultural and Technical College, Hamilton, New Zealand. February, 2003 6. Havemeyer Foundation Workshop on Embryonic and Fetal Nutrition, Ravello, Italy. May, 7. Irish Thoroughbred Breeders’ Association, Co. Kildare, Ireland. October, 2003 8. Havemeyer Foundation Workshop, 3rd Meeting of the European Gamete Group (EEGG), Pardubice, Czech Republic. October, 2003 9. Inner Mongolia University, Hohhot City, China. November, 2003 10. Emerging Equine Science Conference, Royal Agricultural College, Cirencester, UK. 11. Havemeyer Foundation Workshop on Maternal Recognition of Pregnancy in the Mare III, Barbados, West Indies. November, 2004 12. A Workshop on the Equine Placenta, Kentucky, USA. December, 2004 13. Havemeyer Foundation Workshop on Comparative Placentology, Victoria, Canada. April 14. 44th British Equine Veterinary Association Congress, Harrogate, Yorkshire, UK. 15. Havemeyer Foundation Workshop, 1st International Equine Gametes Group Workshop, 16. Thoroughbred Racing and Breeding Seminar, Cheltenham, UK. November, 2005 17. New Zealand Link Foundation and Equibreed Breeders Symposium, Cambridge, New 18. Havemeyer Foundation 2nd Workshop on Embryonic and Fetal Nutrition, Ravello, Italy. 19. Ghent Veterinary School, Ghent, Belgium. March, 2007 20. International Scientist Forum on Mammalian Reproductive Biotechnology, Inner Mongolia University, Hohhot City, Inner Mongolia. August, 2007 21. 5th International Conference on Equine Reproductive Medicine, Leipzig, Germany. 22. Waikato Thoroughbred Breeders’ Association, Cambridge, New Zealand. March, 2007 23. Havemeyer Foundation Workshop, Peter Rossdale 80th Birthday Symposium, Cambridge, 24. Equine Symposium, Alter Real Stud, Portugal. June, 2008 Seminar of Biotechnology and Reproductive Biology, School of Veterinary Medicine, 25. Temuco, Chile. September, 2008 26. 3rd Annual Equine Science Conference, Open College of Equine Studies, Boxted Hall, 27. 27. Rossdale and Partners Equine Pregnancy Failure Veterinary Course, Newmarket, UK. 28. 28. Havemeyer Mini-Symposium on Thoroughbred Breeding Efficiency, Keeneland, Kentucky, 29. University of Georgia, Athens, USA. Embryo Transfer Course. October 2009 30. Havemeyer Mini-Symposium on Thoroughbred Breeding Efficiency, Sydney, Australia. 31. Havemeyer Mini-Symposium on Thoroughbred Breeding Efficiency, Auckland, New 32. Waikato Veterinary Association, Cambridge, New Zealand. March 2010 33. Workshop on Early Embryonic Death, 10th International Symposium on Equine Reproduction, Kentucky USA. July 2010 34. Havemeyer Foundation 30th Anniversary Workshop. Saratoga, USA. August 2011 35. Sharjah Equine Hospital, Sharjah, UAE. Artificial Insemination and Embryo Transfer 36. Newmarket Equine Hospital Advances in Equine Reproduction Course, Newmarket, UK. 37. Colloquium for Equine Reproduction – British Society of Animal Science Annual Meeting 38. Zimbabwe Veterinary Association, Harare, Zimbabwe. June 2013 39. 52nd British Equine Veterinary Association Congress, Manchester, UK. September, 2013 Submitted abstracts Oral presentations
1. 50th Annual Meeting of the European Association Animal Production, Basel, Switzerland. 2. Society for Study of Reproduction and Fertility, Edinburgh, Scotland. July, 2000 3. 5th International Symposium on Equine Embryo Transfer, Saari, Finland. July, 2000 4. 9th Conference of the European Placenta Group, Sorrento, Italy. September, 2001 5. 8th International Symposium on Equine Reproduction, Colorado, USA. August, 2002 6. 7th Annual Conference of the European Society for Domestic Animal Reproduction, Dublin, 7. 6th International Symposium on Equine Embryo Transfer, Rio de Janeiro, Brazil. August, 8. 9th Annual Conference of the European Society for Domestic Animal Reproduction, 9. 9th International Symposium on Equine Reproduction, Kerkade, The Netherlands. August, 10. 11th Annual Conference of the European Society for Domestic Animal Reproduction, Celle, 11. 7th International Symposium on Equine Embryo Transfer, Cambridge, UK. July, 2008 12. 25th European Embryo Transfer Association, Poznan, Poland, September, 2009 13. 10th International Symposium on Equine Reproduction, Kentucky, July 2010 14. 8th International Symposium on Equine Embryo Transfer, Canada. July, 2012 Poster presentations
1. 5th Annual Conference of the European Society for Domestic Animal Reproduction, 2. 8th Meeting of the International Federation of Placenta Associations, Melbourne, 3. 15th International Conference on Animal Reproduction, Porto Segura, Brazil. August, 2004 4. 10th Annual Conference of the European Society for Domestic Animal Reproduction, Ljubljana, Slovenia. September, 2006 5. Fertility 2007: Biennial Meeting of the UK Fertility Societies, York, UK. July, 2007 6. 25th European Embryo Transfer Association, Poznan, Poland 2009 7. 26th European Embryo Transfer Association, Uppsala, Sweden 2010

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